Raised on 1 September 1938 at Reading, Berkshire, to command all the Territorial Army Anti-Aircraft units in the South, South West and South Midlands of England and South Wales.
When Anti-Aircraft Command was re-organised in October 1942, the division was amalgamated into 3 Anti-Aircraft Group.
HQ Reading, Berkshire
Order of Battle 1939
| 35 Anti-Aircraft Brigade |
| 56 (Cornwall) Anti-Aircraft Regiment – unit converted from medium artillery in 1932 |
| 57 (Wessex) Anti-Aircraft Regiment – unit converted from field artillery in 1932 |
| 72 (Hampshire) Anti-Aircraft Regiment – unit converted from field artillery (originally Yeomanry cavalry) in 1938 |
| 48 (Hampshire) Anti-Aircraft Battalion, Royal Engineers – Searchlight unit converted from fortress engineers in 1937 |
| 45 Anti-Aircraft Brigade – Transfer not complete until 1 May 1940 |
| 77 (Welsh) Anti-Aircraft Regiment – unit converted from field artillery in 1938 |
| 6 (Glamorgan) Battalion, Welch Regiment (67 Searchlight Regiment) – infantry battalion converted to searchlight in 1938 |
| 1st (Rifle) Battalion, Monmouthshire Regiment (68 Searchlight Regiment) – infantry battalion converted in 1938 |
| No 2 (Anti-Aircraft Searchlight) Company, Carmarthenshire Fortress Royal Engineers |
| 46 Anti-Aircraft Brigade |
| 76 (Gloucestershire) Anti-Aircraft Regiment – unit converted from field artillery in 1938 |
| 98 Anti-Aircraft Regiment – new unit formed in 1939 |
| 4th (City of Bristol) Battalion, Gloucestershire Regiment (66 Searchlight Regiment) – infantry battalion converted in 1938 |
| 47 Anti-Aircraft Brigade |
| 80 (Berkshire) Anti-Aircraft Regiment – unit converted from field artillery in 1938 |
| 30 (Surrey) Anti-Aircraft Battalion, Royal Engineers – searchlight battalion formed in 1924 |
| 35 (First Surrey Rifles) Anti-Aircraft Battalion, Royal Engineers – infantry battalion converted in 1935 |
| 4th Battalion Queen’s Royal Regiment (West Surrey) (63 Searchlight Regiment) – infantry battalion converted in 1938 |
| 72 (Middlesex) Searchlight Regiment – new unit formed in 1938 |
| 55 Light Anti-Aircraft Brigade |
| 23 Light Anti-Aircraft Regiment – new unit formed in 1938 |
| 24 Light Anti-Aircraft Regiment – new unit formed in 1938 |
| 34 Light Anti-Aircraft Regiment – new unit formed in 1939 |
| 35 Light Anti-Aircraft Regiment – new unit formed in 1939 |
| 36 Light Anti-Aircraft Regiment – new unit formed in 1939 |
Order of Battle 1940
| 5 Anti-Aircraft Brigade – at Gloucester – to 9 Anti-Aircraft Division Nov 40 |
| 84 Heavy Anti-Aircraft Regiment |
| 85 (Tees) Heavy Anti-Aircraft Regiment |
| 47 Light Anti-Aircraft Regiment |
| 37 (Tyne Electrical Engineers) Searchlight Regiment |
| 68 Searchlight Regiment |
| 27 (Home Counties) Anti-Aircraft Brigade – at Portsmouth – from the 6 Anti-Aircraft Division |
| 31 (City of London Rifles) Searchlight Regiment |
| 70 (Sussex) Searchlight Regiment |
| 35 Anti-Aircraft Brigade – at Portsmouth |
| 72 (Hampshire) Heavy Anti-Aircraft Regiment |
| 80 (Berkshire) Heavy Anti-Aircraft Regiment |
| 118 Heavy Anti-Aircraft Regiment – new unit formed Dec 40 |
| 48 (Hampshire) Searchlight Regiment |
| 45 Anti-Aircraft Brigade – Nov 40 to 9 Anti-Aircraft Division |
| 77 (Welsh) Heavy Anti-Aircraft Regiment |
| 79 (Herts Yeomanry) Heavy Anti-Aircraft Regiment |
| 85 (Tees) Heavy Anti-Aircraft Regiment |
| 88 Heavy Anti-Aircraft Regiment |
| 20 Light Anti-Aircraft Regiment |
| 34 Light Anti-Aircraft Regiment |
| 67 (Welch) Searchlight Regiment |
| 77 Searchlight Regiment |
| 47 Anti-Aircraft Brigade – at Southampton |
| 27 (London Electrical Engineers) Searchlight Regiment |
| 30 (Surrey) Searchlight Regiment |
| 63 (Queen’s) Searchlight Regiment |
| 65 Anti-Aircraft Brigade – at Southampton – new formation |
| 57 (Wessex) Heavy Anti-Aircraft Regiment |
| 24 Light Anti-Aircraft Regiment |
| 43 Light Anti-Aircraft Regiment – new unit formed Nov 39 |
| 58 (Argyll & Sutherland Highlanders) Light Anti-Aircraft Regiment – from British Expeditionary Force |
| Divisional unit |
| 5 Anti-Aircraft Z Regiment – formed in Sep 40 |
Order of Battle 1941-42
| 5 Anti-Aircraft Brigade – joined from the 9 Anti-Aircraft Division June 1942 |
| 64 (Northumbrian) Heavy Anti-Aircraft Regiment – from 72 Anti-Aircraft Brigade Aug 42 |
| 97 (London Scottish) Heavy Anti-Aircraft Regiment – from 35 Anti-Aircraft Brigade Jun 42; left Anti-Aircraft Command Aug 42, later went to Sicily and Italy |
| 106 Heavy Anti-Aircraft Regiment – joined from 2 Anti-Aircraft Division Jul 42 |
| 128 Heavy Anti-Aircraft Regiment – joined from the 10 Anti-Aircraft Division Jun; to 2 Anti-Aircraft Division Jul 42 |
| 143 (Mixed) Heavy Anti-Aircraft Regiment – to 11 Anti-Aircraft Division Jun 42 |
| 148 (Mixed) Heavy Anti-Aircraft Regiment – from 35 Anti-Aircraft Brigade Jun 42 |
| 46 Light Anti-Aircraft Regiment – from 65 Anti-Aircraft Brigade Jun 42 |
| 84 Light Anti-Aircraft Regiment – from 12 Anti-Aircraft Division Aug 42 |
| 97 Light Anti-Aircraft Regiment – from 65 Anti-Aircraft Brigade Jun 42 |
| 112 Light Anti-Aircraft Regiment |
| 37 Searchlight Regiment – to 11 Anti-Aircraft Division Jun 42 |
| 27 Anti-Aircraft Brigade |
| 107 Heavy Anti-Aircraft Regiment – joined from 4 Anti-Aircraft Division Jun 41; to 35 Anti-Aircraft Brigade Jun 42 |
| 146 Heavy Anti-Aircraft Regiment – joined from 7 Anti-Aircraft Division Aug 42 |
| 124 Heavy Anti-Aircraft Regiment – from 35 Anti-Aircraft Brigade Autumn 41 |
| 68 Light Anti-Aircraft Regiment – joined from 7 Anti-Aircraft Division Aug 42 |
| 98 Light Anti-Aircraft Regiment – from 72 Anti-Aircraft Brigade Aug 42 |
| 132 Light Anti-Aircraft Regiment – formed from 85 Searchlight Regiment Mar 42; to 6 Anti-Aircraft Division Jun 42 |
| 1 Searchlight Regiment – joined from 9 Anti-Aircraft Division Jan 42 |
| 31 Searchlight Regiment |
| 34 Searchlight Regiment |
| 35 Searchlight Regiment – returned summer 41; to 47 Anti-Aircraft Brigade Dec 41 |
| 35 Anti-Aircraft Brigade |
| 54 (City of London) Heavy Anti-Aircraft Regiment – joined from 1 Anti-Aircraft Division Autumn 41; returned to 1 Anti-Aircraft Division Feb 42 |
| 57 Heavy Anti-Aircraft Regiment – left for 1 Anti-Aircraft Division Autumn 41 |
| 72 Heavy Anti-Aircraft Regiment – left for 6 Anti-Aircraft Division Dec 41 |
| 80 Heavy Anti-Aircraft Regiment – left Anti-Aircraft Command Jul 41; later went to North Africa |
| 97 (London Scottish) Heavy Anti-Aircraft Regiment – joined from 1 Anti-Aircraft Division Mar 42; to 5 Anti-Aircraft Brigade Jun 42 |
| 101 Heavy Anti-Aircraft Regiment – joined from 6 Anti-Aircraft Division Jul 42; left for 1 Anti-Aircraft Division Aug 42 |
| 104 Heavy Anti-Aircraft Regiment – joined from 8 Anti-Aircraft Division Dec 41, left for 6 Anti-Aircraft Division Apr 42 |
| 107 Heavy Anti-Aircraft Regiment – from 27 Anti-Aircraft Brigade Jun 42 |
| 124 Heavy Anti-Aircraft Regiment – new unit formed Mar 41 to 27 Anti-Aircraft Brigade Autumn 41 |
| 148 (Mixed) Heavy Anti-Aircraft Regiment – new unit formed Feb 42 to 5 Anti-Aircraft Brigade Jun 42 |
| 151 (Mixed) Heavy Anti-Aircraft Regiment – from 4 Anti-Aircraft Division Jul 42 |
| 157 (Mixed) Heavy Anti-Aircraft Regiment – new unit formed May 42 |
| 160 (Mixed) Heavy Anti-Aircraft Regiment – new unit formed Jun 42 |
| 48 Searchlight Regiment – from 47 Anti-Aircraft Brigade Aug 42 |
| 5 Anti-Aircraft Z Regiment |
| 47 Anti-Aircraft Brigade |
| 35 Searchlight Regiment – from 27 Anti-Aircraft Brigade Dec 41 |
| 48 Searchlight Regiment – to 35 Anti-Aircraft Brigade Aug 42 |
| 63 Searchlight Regiment (Queens) – left for 1 Anti-Aircraft Division Autumn 41 |
| 70 Searchlight Regiment |
| 65 Anti-Aircraft Brigade – left for the 10 Anti-Aircraft Division May 1942 |
| 23 Light Anti-Aircraft Regiment – left Anti-Aircraft Command Dec 41, later went to Ceylon |
| 24 Light Anti-Aircraft Regiment – left for 8 Anti-Aircraft Division Summer 41 |
| 43 Light Anti-Aircraft Regiment – left for 6 Anti-Aircraft Division Jun 41 |
| 46 Light Anti-Aircraft Regiment – from 55 Anti-Aircraft Brigade Summer 41 |
| 60 Light Anti-Aircraft Regiment – joined from 12 Anti-Aircraft Division Autumn 41 |
| 80 Light Anti-Aircraft Regiment – new unit formed Aug 41; left for 9 Anti-Aircraft Division Dec 41 |
| 81 Light Anti-Aircraft Regiment – joined from 6 Anti-Aircraft Division Mar 42 |
| 97 Light Anti-Aircraft Regiment – new unit formed Nov 41; to 5 Anti-Aircraft Brigade Jun 42 |
| 72 Anti-Aircraft Brigade – new formation, joined June 1942 |
| 64 (Northumbrian) Heavy Anti-Aircraft Regiment – joined Jun 42 ; to 5 Anti-Aircraft Brigade Aug 42 |
| 157 (Mixed) Heavy Anti-Aircraft Regiment – joined Jun 42 |
| 110 Light Anti-Aircraft Regiment – joined Aug 42 |
| 123 Light Anti-Aircraft Regiment – joined Aug 42 |
| 50 (Northamptonshire Regiment) Searchlight Regiment – joined Jun 42 |